The Freshwater Diatoms of Ireland

2011 Richard T. Carter


This revision of the Irish checklist by Richard T. Carter incorporates many taxonomic changes that have been made in the literature since the first version. It must not be regarded as the last word on the Irish fresh-water diatom flora, however, as almost every new sample that we look at contains additional taxa. There are also numerous taxa that remain to be identified. As in the previous version, the list incorporates floral lists from all the publications on Irish freshwater diatoms to which he has access. Also, as in the previous version, currently accepted names are left-justified, while common synonyms in the Irish literature are indented beneath the currently accepted name.


A word of caution is definitely in order: it is almost always the case that, in long-established diatom genera, one or more specific epithets have been used as "catch-all" names for groups of closely-related taxa. In the genus Stauroneis, for example, there are three such names: S. phoenicenteron (Nitzsch) Ehrenberg, S. gracilis Ehrenberg, and S. anceps Ehrenberg. When we see one of these names on an older floral list, we have very little idea as to which (if any, since new species are discovered with every floral study) of the currently accepted taxa the author may have had. When one sees S. phoenicenteron on an older floral list, it normally means only "a big Stauroneis species". (The group of species around S. anceps has grown particularly large, making this name particularly suspect when examining older floras.) The situation in other large and long-established genera is the same, and it is wise to learn which specific epithets are the "catch-alls" for their genus.


Valid taxa are justified left; synonyms are right-indented. 

Preparation of this list was greatly facilitated through the continual use of:

Catalogue of Diatom Names, California Academy of Sciences, On-line Version.

Compiled by Elisabeth Fourtanier & J. Patrick Kociolek.

Available online at


Additions and corrections to this checklist will be gratefully accepted.


Freshwater genera A-E


ACANTHOCERAS Honigmann 1910

 A. zachariasii (Brun) Simonsen

Attheya zachariasii Brun 



A. altaica(Poretzky) Cleve-Euler

A. borealis A. Cleve

A. brevipes Agardh

A. brevipes var. intermedia (Kützing) Cleve

A. brevipes var. parvula (Kützing) Cleve

A. coarctata (Brébisson) Grunow

A. detha Hohn & Hellerman

A. gysingensis (Foged) Lange-Bertalot in Lange-Bertalot & Krammer

Navicula gysingensis Foged

A. impexa Lange-Bertalot in Lange-Bertalot & Krammer

A. obliqua (Gregory) Hustedt

A. recurvata Hustedt

A. rostellata Cleve-Euler

A. saxonica Krasske in Hustedt

A. ventralis (Krasske) Lange-Bertalot in Lange-Bertalot & Krammer

Navicula ventralis Krasske 



 A. biasolettianum (Kützing) Bukhtiyarova

Achnanthes biasolettiana (Kützing) Grunow

A. cryptocephalum (Grunow in Van Heurck) Mills

Achnanthes minutissima var. cryptocephala Grunow in Van Heurck

A. exiguum (Grunow) Czarnecki

Achnanthes exigua Grunow

A. exiguum var. heterovalvatum (Krasske) Czarnecki

Achnanthes exigua var. heterovalvata Krasske

A. exile (Kützing) Heiberg

Achnanthes exilis Kützing

A. helveticum (Hustedt) Monnier, Lange-Bertalot & Ector in Monnier et al.

Achnanthes helvetica (Hustedt) Lange-Bertalot in Lange-Bertalot & Krammer

A. minutissimum (Kützing) Czarnecki

Achnanthes minutissima Kützing

A. minutissimum var. affinis (Grunow) Bukhtiyarova

Achnanthes affinis Grunow

A. subatomoides (Hustedt) Monnier, Lange-Bertalot & Ector in Monnier et al.

Navicula subatomoides Hustedt

A. trinode Ralfs in Pritchard

Achnanthes trinodis (Ralfs in Pritchard) Grunow in Van Heurck  


ADLAFIA Moser, Lange-Bertalot & Metzeltin 1998

 A. bryophila (Petersen) Moser, Lange-Bertalot & Metzeltin

Navicula bryophila Petersen

A. minuscula (Grunow) Lange-Bertalot in Lange-Bertalot & Genkal

Navicula minuscula Grunow in Van Heurck

A. minuscula var. muralis (Grunow) Lange-Bertalot in Lange-Bertalot & Genkal

Navicula muralis Grunow in Van Heurck

A. suchlandtii (Hustedt) Moser, Lange-Bertalot & Metzeltin

Navicula suchlandtii Hustedt 


 AMPHIPLEURA Kützing 1844

 A. pellucida (Kützing) Kützing 


AMPHORA Ehrenberg ex Kützing 1844

 A. copulata (Kützing) Schoeman & Archibald

A. ovalis var. libyca (Ehrenberg) Cleve

A. libyca Ehrenberg

A. inariensis Krammer

A. ovalis (Kützing) Kützing

A. pediculus (Kützing) Grunow ex Schmidt

A. ovalis var. pediculus (Kützing) Van Heurck

A. parallelistriata Manguin ex Kociolek & Reviers

A. perpusilla (Grunow in Van Heurck) Grunow in Van Heurck


ANEUMASTUS D. Mann & Stickle in Round et al. 1990

A. minor (Hustedt) Lange-Bertalot

A. tuscula f. minor (Hustedt) Bukhtiyarova

Navicula tuscula f. minor Hustedt

A. rostrata (Hustedt) Lange-Bertalot

A. tuscula f. rostrata (Hustedt) Bukhtiyarova

Navicula tuscula f. rostrata Hustedt

A. stroesei (Østrup) D. Mann in Round et. al.

Navicula pseudotuscula Hustedt

A. tuscula (Ehrenberg) D. Mann & Stickle in Round et al.

Navicula tuscula Ehrenberg

A. tusculoides (Cleve-Euler) D. Mann in Round et al.

Navicula tusculoides Cleve-Euler 


ANOMOEONEIS  Pfitzer 1871

 A. sphaerophora (Kützing) Pfitzer 



 A. formosa Hassall

A. gracillima (Hantzsch in Rabenhorst) Heiberg

A. ralfsii W. Smith 


AULACOSEIRA Thwaites 1848

 A. alpigena (Grunow) Krammer

Melosira distans var. alpigena Grunow in Van Heurck

 A. ambigua (Grunow) Simonsen

Melosira ambigua (Grunow in Van Heurck) O.Müller

A. crenulata (Ehrenberg) Krammer

A. distans (Ehrenberg) Simonsen

Melosira distans (Ehrenberg) Kützing

A. fennoscandica (A. Cleve-Euler) Ross in Hartley

Melosira fennoscandica Cleve-Euler

A. granulata (Ehrenberg) Simonsen

Melosira granulata (Ehrenberg) Ralfs in Pritchard

A. granulata var. angustissima (O. Müller) Simonsen

Melosira granulata var. angustissima O. Müller

A. granulata f. curvata (Hustedt) Simonsen

Melosira granulata f. curvata Hustedt

A. islandica (O. Müller) Simonsen

Melosira islandica O. Müller

A. islandica ssp. helvetica (O. Müller) Simonsen

Melosira islandica ssp. helvetica O. Müller

A. italica (Ehrenberg) Simonsen

Melosira italica (Ehrenberg) Kützing

A. lacustris (Grunow in Van Heurck) Krammer

Melosira distans f. lacustris (Grunow in Van Heurck) Hustedt

? Melosira mikkelsenii Nygaard

A. lirata (Ehrenberg) Ross

Melosira distans var. lirata (Ehrenberg) O. Müller

A. lirata var. seriata (O. Müller) Vyverman

Melosira distans f. seriata (O. Müller) Hustedt

A. perglabra (Østrup) Haworth

? Melosira excurrens Nygaard

A. subarctica (O. Müller) Haworth

Melosira italica ssp. subarctica O. Müller

A. valida (Grunow) Krammer

Melosira italica var. valida Grunow 


BERKELEYA Greville 1827

B. rutilans (Trentepohl ex Roth) Grunow

Amphipleura rutilans (Trentepohl ex Roth) Cleve 


BIREMIS D. Mann & Cox in Round et al. 1990

B. zachariasii (Reichelt) Edlund, Andresen & Soninkhisig

Oestrupia zachariasii (Reichelt) Hustedt 


 BRACHYSIRA Kützing 1836

 B. brebissonii Ross in Hartley

Anomoeoneis serians var. brachysira (Brébisson) Cleve

Anomoeoneis serians var. thermalis (Grunow) Hustedt

B. exilis (Kützing) Round & Mann

Anomoeoneis exilis (Kützing) Cleve

B. follis (Ehrenberg) Ross in Hartley

Anomoeoneis follis (Ehrenberg) Cleve

B. microcephala (Kützing) Compère

Achnanthes microcephala (Kützing) Grunow

B. serians (Brébisson) Round & Mann

Anomoeoneis serians (Brébisson) Cleve

B. styriaca (Grunow in Van Heurck) Ross in Hartley

Anomoeoneis styriaca (Grunow in Van Heurck) Hustedt

B. vitrea (Grunow) Ross in Hartley

Anomoeoneis vitrea (Grunow) Ross in Patrick & Reimer

B. vitrea f. lanceolata (Mayer) Schoeman &  Archibald

Anomoeoneis exilis f. lanceolata (A. Mayer) Hustedt

B. zellensis (Grunow) Round & Mann

Anomoeoneis zellensis (Grunow) Cleve

B. zellensis f. difficilis (Grunow in Van Heurck) Hamilton

Anomoeoneis zellensis f. difficilis (Grunow) Hustedt 


CALONEIS Cleve 1894

C. alpestris (Grunow) Cleve

C. amphisbaena (Bory) Cleve

C. bacillaris (Gregory) Cleve

C. bacillum (Grunow) Cleve

Pinnularia fasciata (Lagerstedt) Hustedt

C. clevei (Lagerstedt) Cleve

C. formosa (Gregory) Cleve

C. hebes (Ralfs in Pritchard) Patrick

C. obtusa (W. Smith) Cleve

C. latiuscula (Kützing) Cleve

C. latiuscula var. subholstei Hustedt

C. leptosoma (Grunow) Krammer in Krammer & Lange-Bertalot

Pinnularia leptosoma (Grunow in Van Heurck) Cleve

C. limosa (Kützing) Patrick

C. schumanniana (Grunow) Cleve

C. silicula var. gibberula (Kützing) Grunow

C. limosa var. biconstricta (Grunow) Foged

C. schumanniana var. biconstricta (Grunow) Reichelt

C. macedonica Hustedt

C. molaris (Grunow) Krammer in Krammer & Lange-Bertalot

Pinnularia molaris (Grunow) Cleve

C. obtusa (W. Smith) Cleve

C. schumanniana f. fontinalis Foged1

C. sublinearis (Grunow) Krammer in Krammer & Lange-Bertalot

Pinnularia sublinearis (Grunow in Van Heurck) Cleve

C. undulata (Gregory) Krammer in Krammer & Lange-Bertalot

Pinnularia undulata Gregory

? Pinnularia undulata var. stauroneiformis (Hustedt) Cleve-Euler

C. ventricosa (Ehrenberg) Meister

C. silicula (Ehrenberg) Cleve

C. ventricosa var. truncatula (Grunow) Meister

C. silicula var. truncatula (Grunow) Cleve 


CAMPYLODISCUS Ehrenberg ex Kützing 1844

C. hibernicus Ehrenberg

C. noricus Ehrenberg 


CAVINULA D. Mann & Stickle in Round et. al. 1990

C. cocconeiformis (Gregory ex Greville) D. Mann & Stickle

Navicula cocconeiformis Gregory ex Greville

C. jaernefeltii (Hustedt) D. Mann & Stickle

Navicula jaernefeltii Hustedt

C. lacustris (Gregory) D. Mann & Stickle

Navicula lacustris Gregory

C. lapidosa (Krasske) Lange-Bertalot in Lange-Bertalot & Metzeltin

Navicula lapidosa Krasske

C. pseudoscutiformis (Hustedt) D. Mann & Stickle

Navicula pseudoscutiformis Hustedt

C. scutelloides (W. Smith) Lange-Bertalot in Lange-Bertalot & Metzeltin

Navicula scutelloides W. Smith

C. variostriata (Krasske) D. Mann in Round et al.

Navicula variostriata Krasske 


CHAMAEPINNULARIA Lange-Bertalot & Krammer in Lange-Bertalot & Metzeltin 1996

C. krookii (Grunow) Lange-Bertalot & Krammer in Lange-Bertalot & Genkal

Pinnularia krookei Grunow  (Foged, sic)

C. mediocris (Krasske) Lange-Bertalot in Lange-Bertalot & Metzeltin

Navicula mediocris Krasske

C. soehrensis var. hassiaca (Krasske) Lange-Bertalot in Lange-Bertalot & Metzeltin

Navicula hassiaca Krasske 


COCCONEIS Ehrenberg 1837


C. disculus (Schumann) Cleve

C. disculus var. diminuta (Pantocsek) Cleve-Euler

C. diminuta Pantocsek

C. hustedtii Krasske

C. pediculus Ehrenberg

C. placentula Ehrenberg

C. placentula var. euglypta (Ehrenberg) Grunow

C. placentula var. klinoraphis Geitler

C. placentula var. lineata (Ehrenberg) Van Heurck

C. scutellum Ehrenberg

C. scutellum var. parva Grunow in Van Heurck

C. thumensis Mayer 


COSMIONEIS D. Mann & Stickle in Round et al. 1990

C. pusilla (W. Smith) D. Mann & Stickle in Round et al.

Navicula pusilla W. Smith 


CRATICULA Grunow 1867

C. accomoda (Hustedt) D. Mann in Round et al.

Navicula accomoda Hustedt

C. ambigua (Ehrenberg) D. Mann in Round, Crawford & Mann

Navicula cuspidata var. ambigua (Ehrenberg) Cleve

C. cuspidata (Kützing) D. Mann in Round, Crawford & Mann2

Navicula cuspidata (Kützing) Kützing

Navicula cuspidata var. heribaudii M. Peragallo in Héribaud

C. halophila (Grunow ex Van Heurck) D. Mann in Round, Crawford & D. Mann

Navicula halophila Grunow in Van Heurck 


CTENOPHORA Williams & Round 1986

C. pulchella (Ralfs ex Kützing) Williams & Round

Fragilaria pulchella (Ralfs ex Kützing) Lange-Bertalot

Synedra pulchella (Ralfs) Kützing

? Synedra pulchella var. naviculacea Grunow in Van Heurck 


CYCLOSTEPHANOS Round in Theriot et al. 1987

C. dubius (Fricke) Round

Stephanodiscus dubius (Fricke) Hustedt

C. invisitatus (Hohn & Hellerman) Theriot, Stoermer & Håkansson

C. tholiformis Stoermer, Håkansson & Theriot 


CYCLOTELLA (Kützing) Brébisson 1838

C. atomus Hustedt

C. comensis Grunow in Van Heurck

C. ocellata Pantocsek

C. kuetzingiana var. planetophora Fricke

C. rossii Håkansson

C. comta var. oligactis (Ehrenberg) Grunow in Van Heurck

C. striata (Kützing) Grunow

C. striata var. bipunctata Fricke

C. unipunctata Håkansson & Carter 



C. elliptica (Brébisson) W. Smith

C. elliptica var. constricta Grunow

C. elliptica var. hibernica (W. Smith) Van Heurck

C. solea (Brébisson) W. Smith

C. librile (Ehrenberg) Pantocsek

C. solea var. graciloides Cleve-Euler

C. solea var. regula (Ehrenberg) Grunow

C. solea var. apiculata (W. Smith) Ralfs in Pritchard

C. librile var. apiculata (W. Smith) Descy


CYMBELLA Agardh 1830

C. affinis Kützing

C. tumidula Grunow in Schmidt

C. aspera (Ehrenberg) Cleve3

C. cistula (Ehrenberg in Hemprich & Ehrenberg) Kirchner4

C. cistula var. maculata (Kützing) Van Heurck

C. compacta Østrup

C. helvetica var. compacta (Østrup) Hustedt

C. cymbiformis Agardh

C. cymbiformis var. subrhomboidea Østrup

C. designata Krammer in Krammer & Lange-Bertalot

C. helvetica Kützing

C. hustedtii Krasske

C. laevis Nägeli in Kützing

C. lanceolata (Ehrenberg) Van Heurck

C. leptoceros (Ehrenberg) Kützing5

C. obtusa Gregory

C. pseudonorvegica Haworth

C. tumida (Brébisson in Kützing) Van Heurck

C. turgidula Grunow in Schmidt 


CYMBELLAFALSA Lange-Bertalot & Metzeltin 2009

C. diluviana (Krasske) Lange-Bertalot & Metzeltin in Metzeltin et al.

Cymbella diluviana (Krasske) Florin



C. diluviana Hustedt 


CYMBOPLEURA Krammer 1999

C. amphicephala (Nägeli) Krammer

Cymbella amphicephala Nägeli ex Kützing

C. angustata (W. Smith) Krammer

Cymbella angustata (W. Smith) Cleve

C. austriaca (Grunow) Krammer

Cymbella austriaca Grunow in Schmidt

C. cuspidata (Kützing) Krammer

Cymbella cuspidata Kützing

? Cymbella obtusiuscula Kützing

C. hauckii (Van Heurck) Krammer

Cymbella hauckii Van Heurck

C. hybrida (Grunow) Krammer

Cymbella hybrida Grunow

C. inaequalis (Ehrenberg) Krammer

Cymbella inaequalis (Ehrenberg) Rabenhorst

Cymbella ehrenbergii Kützing

C. inaequilatera (Lagerstedt) Krammer

Cymbella aequalis W. Smith ex Greville

C. incerta (Grunow)  Krammer

Cymbella incerta Grunow

C. incertiformis var. linearis (Fontell) Krammer

Cymbella angustata var. linearis (Fontell) Ross

C. lata (Grunow) Krammer

Cymbella lata Grunow

C. naviculiformis (Auerswald ex Heiberg) Krammer

Cymbella naviculiformis Auerswald ex Heiberg

C. neoheteropleura Krammer

Cymbella heteropleura (Ehrenberg) Kützing

C. reinhardtii (Grunow) Krammer

C. similis (Krasske) Krammer

Cymbella similis Krasske

C. subcuspidata (Krammer) Krammer

Cymbella subcuspidata Krammer

Cymbella heteropleura var. minor Cleve 

DECUSSATA (Patrick) Lange-Bertalot 2000

D. placenta (Ehrenberg) Lange-Bertalot & Metzeltin

          Navicula placenta Ehrenberg 


DELICATA Krammer 2003

D. delicatula (Kützing) Krammer

          Cymbella delicatula Kützing 


DENTICULA Kützing 1844

D. kuetzingii Grunow

          Nitzschia denticula Grunow in Cleve & Grunow

D. tenuis Kützing

          D. tenuis var. crassula (Nägeli) Hustedt 


DIADESMIS Kützing 1844

D. contenta (Grunow in Van Heurck) D. Mann

          Navicula contenta Grunow in Van Heurck

D. contenta var. biceps (Grunow in Van Heurck) Hamilton in Hamilton et al.

          Navicula contenta var. biceps Grunow in Van Heurck

D. contenta var. parallela (Petersen) Spaulding in Spaulding et al.

          Navicula contenta var. parallela Petersen

D. gallica var. perpusilla (Grunow) Bukhtiyarova

D. perpusilla (Grunow) D. Mann in Round et al.

Navicula perpusilla Grunow 


DIATOMA Bory 1824

D. ehrenbergii Kützing

          D. vulgare var. ehrenbergii (Kützing) Grunow

          D. vulgare var. grande (W. Smith) Grunow

D. hiemale (Lyngbye) Heiberg

D. mesodon (Ehrenberg) Kützing

          D. hiemale var. mesodon (Ehrenberg) Grunow

D. moniliforme Kützing

D. tenue Agardh

          D. elongatum (Lyngbye) Agardh

          D. elongatum var. minor Grunow

          D. elongatum var. tenuis (Agardh) Van Heurck

D. vulgare Bory

D. vulgare var. linearis Grunow in Van Heurck

D. vulgare var. producta Grunow 


DIATOMELLA Greville 1855

D. balfouriana Greville 


DIDYMOSPHENIA  M. Schmidt 1899

D. geminata (Lyngbye) M. Schmidt  


DIPLONEIS  Ehrenberg 1844

D. boldtiana Cleve

D. domblittensis (Grunow) Cleve

D. domblittensis var. subconstricta A. Cleve

D. elliptica (Kützing) Cleve

D. finnica (Ehrenberg) Cleve

D. interrupta (Kützing) Cleve

D. marginestriata Hustedt

D. minuta Petersen

D. oblongella (Nägeli ex Kützing) Ross

D. ovalis var. oblongella (Nägeli) Cleve

D. oculata (Brébisson) Cleve

D. ovalis (Hilse in Rabenhorst) Cleve

D. petersenii Hustedt

D. pseudovalis Hustedt

D. puella (Schumann) Cleve

D. smithii var. pumila (Grunow) Hustedt6

D. subovalis Cleve 


DISCOSTELLA Houk & Klee 2004

 D. glomerata (Bachmann) Houk & Klee

          Cyclotella glomerata Bachmann

D. pseudostelligera (Hustedt) Houk & Klee

          Cyclotella pseudostelligera Hustedt

D. stelligera (Cleve & Grunow) Houk & Klee

          Cyclotella stelligera Cleve & Grunow in Cleve

D. stelligeroides (Hustedt) Houk & Klee

Cyclotella stelligeroides Hustedt 


ELLERBECKIA Crawford 1988


E. arenaria (Moore ex Ralfs) Crawford

          Melosira arenaria Moore ex Ralfs

E. arenaria f. teres (Brun) Crawford 


ENCYONEMA Kützing 1833

E. auerswaldii Rabenhorst

Cymbella ventricosa var. auerswaldii (Rabenhorst) Meister

E. caespitosum Kützing

E. caespitosum var. semicirculare (Lagerstedt) Mayer

Cymbella ventricosa var. semicircularis (Lagerstedt) Ross

E. elginense(Krammer) D. Mann in Round et al.

Cymbella elginensis Krammer

E. turgidum (Gregory) Grunow in Schmidt

Cymbella turgida Gregory

E. gaeumannii (Meister) Krammer

Cymbella gaeumannii Meister

E. gracile Rabenhorst

Cymbella gracilis (Rabenhorst) Cleve

E. hebridicum (Gregory) Grunow in Cleve & Möller

Cymbella hebridica (Grunow in Cleve & Möller) Cleve

E. hilliardii (Manguin) Krammer

Cymbella hilliardii Manguin

E. lacustre (Agardh) D. Mann

Cymbella lacustris (Agardh) Cleve

E. minutum (Hilse in Rabenhorst) D. Mann

Cymbella minuta Hilse in Rabenhorst

Cymbella ventricosa var. minuta (Hilse in Rabenhorst) Cleve-Euler

E. minutum var. pseudogracilis (Cholnoky) Czarnecki

Cymbella turgida var. pseudogracilis Cholnoky

E. norvegica (Grunow in Schmidt) Bukhtiyarova

Cymbella norvegica Grunow in Schmidt

E. perpusillum (Cleve) D. Mann

Cymbella perpusilla Cleve

E. prostratum (Berkeley) Kützing

Cymbella prostrata (Berkeley) Cleve

E. silesiacum (Bleisch in Rabenhorst) D. Mann in Round et al.

Cymbella silesiaca Bleisch in Rabenhorst

E. ventricosum (Kützing) Grunow in Schmidt

Cymbella ventricosa Kützing 


ENCYONOPSIS Krammer 1997

E. cesatii (Rabenhorst) Krammer

Cymbella cesatii (Rabenhorst) Grunow in Schmidt

E. descripta (Hustedt) Krammer

Cymbella descripta (Hustedt) Krammer & Lange-Bertalot

E. microcephala (Grunow) Krammer

Cymbella microcephala Grunow 


ENTOMONEIS Ehrenberg 1845

E. alata Ehrenberg

Amphiprora alata (Ehrenberg) Kützing

E. ornata (Bailey) Reimer in Patrick & Reimer

Amphiprora ornata Bailey 


EOLIMNA Lange-Bertalot & Schiller in Schiller & Lange-Bertalot 1997

E. minima (Grunow in Van Heurck) Lange-Bertalot in Moser et al.

Navicula minima Grunow

E. rotunda (Hustedt) Lange-Bertalot, Kulikovskiy & Witkowski in Kulikovskiy et al.

Navicula rotunda Hustedt

E. subminuscula (Manguin) Moser, Lange-Bertalot & Metzeltin

Navicula subminuscula Manguin

E. tantula (Hustedt) Lange-Bertalot in Werum & Lange-Bertalot

Navicula tantula Hustedt 


EPITHEMIA Brébisson ex Kützing 1844

E. adnata (Kützing) Brébisson

E. zebra (Ehrenberg) Kützing

E. zebra var. porcellus (Kützing) Grunow

E. zebra var. saxonica (Kützing) Grunow

E. argus (Ehrenberg) Kützing

E. longicornis (Ehrenberg) W. Smith

E. argus var. alpestris (W. Smith) Grunow

E. frickei Krammer

E. intermedia Fricke in Schmidt

E. goeppertiana Hilse

E. muelleri Fricke in Schmidt

E. hyndmannii W. Smith

E. ocellata (Ehrenberg) Kützing7

E. reichelti Fricke

E. smithii Carruthers

E. sorex Kützing

E. sorex var. gracilis Hustedt

E. turgida (Ehrenberg) Kützing

E. turgida var. capitata Fricke

E. turgida var. granulata (Ehrenberg) Brun         


EUCOCCONEIS Cleve ex Meister 1912

E. austriaca (Hustedt) Lange-Bertalot

Achnanthes austriaca Hustedt

E. depressa (Cleve) Lange-Bertalot

Achnanthes depressa (Cleve) Hustedt

E. flexella (Kützing) Meister

Achnanthes flexella (Kützing) Brun

E. flexella var. alpestris (Brun) Hustedt

Achnanthes flexella var. alpestris Brun

E. laevis (Østrup) Lange-Bertalot

Achnanthes laevis Østrup

E. lapponica Hustedt

Achnanthes lapponica Hustedt 


EUNOTIA Ehrenberg 1837

E. arcuoides Foged

E. arcus Ehrenberg

E. arcus var. bidens Grunow in Van Heurck

E. arcus var. fallax Hustedt

E. bactriana Ehrenberg

E. bidentula W. Smith

E. bidentuloides Foged

E. bidens Ehrenberg

E. praerupta var. bidens Grunow in Cleve & Grunow

E. bigibba Kützing sensu Grunow

E. bigibba var. pumila Grunow

E. bilunaris (Ehrenberg) Mills

E. lunaris (Ehrenberg) Grunow in Van Heurck

E. compacta (Hustedt) Mayama

E. exigua var. compacta Hustedt

E. curtagrunowii Nörpel-Schempp & Lange-Bertalot

E. praerupta var. curta Grunow in Van Heurck

E. praerupta var. muscicola Petersen

E. denticulata (Brébisson) Rabenhorst

E. diadema Ehrenberg

E. robusta var. diadema (Ehrenberg) Ralfs in Pritchard

E. diodon Ehrenberg

E. elegans Østrup

E. exigua (Brébisson ex Kützing) Rabenhorst

E. faba Ehrenberg

E. faba f. rhomboidea Foged

E. fallax Cleve8

E. flexuosa (Brébisson) Kützing

E. flexuosa var. pachycephala Grunow in Van Heurck

E. formica Ehrenberg

E. glacialis Meister

E. hemicyclus (Ehrenberg) Ralfs in Pritchard

Amphicampa hemicyclus (Ehrenberg) Karsten

Semiorbis hemicyclus (Ehrenberg) Patrick in Patrick & Reimer

E. iatriaensis Foged

E. incisa Gregory

E. jemtlandica (Fontell) Cleve-Euler

E. monodon var. bidens (Gregory) Hustedt

E. lapponica Grunow ex A. Cleve

E. maior (W. Smith) Rabenhorst9

E. meisteri Hustedt

E. microcephala Krasske

E. microcephala var. tridentata (Mayer) Hustedt

E. minor (Kützing) Grunow in Van Heurck

E. pectinalis var. minor (Kützing) Rabenhorst

E. monodon Ehrenberg

? E. maior var. ventricosa A. Cleve

? E. monodon var. maior (W. Smith) Hustedt

? E. maior var. compacta Berg

? E. maior var. scandia Cleve-Euler

E. naegelii Migula in Thomé

E. alpina (Nägeli) Hustedt

E. papilio (Ehrenberg) Hustedt10

E. parallela Ehrenberg

E. parallela var. angusta Grunow

E. parallela var. densestriata Fontell

E. pectinalis (Dillwyn) Rabenhorst

E. pectinalis var. undulata (Ralfs) Rabenhorst

E. pectinalis var. ventralis (Ehrenberg) Hustedt

E. pectinalis var. ventricosa (Ehrenberg) Grunow in Van Heurck

E. polydentula (Brun) Hustedt

E. praerupta Ehrenberg

E. rhomboidea Hustedt

E. septentrionalis Østrup11

E. serra Ehrenberg

E. robusta Ralfs in Pritchard

E. subarcuatoides Alles, Nörpel & Lange-Bertalot

E. lunaris var. subarcuata (Nägeli) Grunow in Van Heurck

E. sudetica O. Müller

E. sudetica var. bidens Hustedt in Schmidt

E. tenella (Grunow) Hustedt in Schmidt

E. tetraodon Ehrenberg

E. robusta var. tetraodon (Ehrenberg) Ralfs in Pritchard

E. trinacria Krasske

E. valida Hustedt 1930

E. gracilis (Ehrenberg) Rabenhorst

E. variundulata Nörpel-Schempp & Lange-Bertalot in Lange-Bertalot et al.

E. exigua var. bidens Hustedt

E. veneris (Kützing) De Toni 




1.  Since Caloneis schumanniana is a junior synonym of C. limosa, this form should be moved accordingly.

2. Navicula cuspidata var. heribaudii M. Peragallo in Héribaud is the initial cell of C. cuspidata.

3. Cymbella aspera (Ehrenberg) Peragallo is one of the "catch-all" names in the genus, and records for this taxon would need to be reexamined to bring them in line with current taxonomy. Krammer (2002: 114) has this to say: "The identification "C. aspera" relates in the literature up to date to a larger number of taxa, distinguished by outline, fine structure, size and other characters."

4. The name "Cymbella cistula" is another catch-all in this genus. Krammer abandons the name, suggesting that the taxon to which it originally referred is not identifiable. In his opinion, recent records referred to this name represent three distinct species: C. subcistula Krammer, C. neocistula Krammer, and C. dorsenotata Østrup.

5. Krammer has abandoned this name, as the original referent is not identifiable (and probably represented a species of Encyonema). Recent records refer to C. subleptoceros Krammer and C. neoleptoceros Krammer.

6.  Foged (1977) claims that this taxon is synonymous with D. puella (Schumann) Cleve.  This is in error, and it is thus an open question as to which taxon Foged had.

7.  Krammer and Lange-Bertalot (1997) argue that Ehrenberg’s Eunotia ocellata is not recognisable.  They place it with Epithemia argus.

8.  Several new species near E. fallax have been recently described, and Irish specimens in this group need to be re-examined.

9.  The situation as regards E. maior and E. monodon, with their many varieties, is a taxonomic morass.  Whether these taxa are distinct or not, and if so, which varieties go with which species, are questions requiring additional research.

10.  This is almost certainly a misidentification. According to Krammer and Lange-Bertalot (2004), E. papilio is a tropical species, not found in Europe. Perhaps this represents E. pseudopapilio Lange-Bertalot & Nörpel-Schempp in Metzeltin & Lange-Bertalot.


11. It is likely that some specimens identified as E. septentrionalis should be referred instead to the very similar E. ursamaioris Lange-Bertalot & Nörpel-Schempp in Lange-Bertalot & Genkal. Both probably occur in Ireland.